This large hippopotamus is a late representative of the characteristic faunas of the age known as the upper Villafranchiano of the upper Valdarno (dating back to about 2-1.5 million years ago), when the landscape was characterized by open forest environments alternating with savannah environments, in following the drying up and lowering of temperatures at the beginning of the Pleistocene (about 2.6 million years ago). Like the current hippos, it was a large partially aquatic herbivore, whose males were endowed with particularly long tusks, used for defense and for fighting for the territory. It is possible that the ancient hippopotamus even survived up to about 126,000 years ago, at the beginning of the Upper Pleistocene, although the last fossils certainly belonging to this species date back to about 500,000 years ago.